Liverpool Skylilne

School of Biological and Environmental Sciences

Dr Barbara Fruth

Dr Barbara Fruth

Telephone: 0151 231 2147



Reader/Associate Professor in Primate Behaviour and Conservation

I am interested in the evolution of animal social systems and the complex interactions between plants and animals. The Central Congo Basin offers a unique field to explore both topics: split by the Congo River, a geographical barrier separating our closest living relatives, the chimpanzee (Pan troglodytes) and the bonobo (Pan paniscus), its diversity of fauna and flora is extremely rich and still largely unexplored.

Since 1990, I am studying wild bonobos in the Democratic Republic of the Congo, a species that has evolved striking peculiarities being contrary to biological paradigms, as well as to its sister species, the chimpanzee. Apart from bonobo social behaviour, their ecological constraints and role within the ecosystem, I am specifically interested in their health status as a direct measure of fitness. In this context I am investigating the transition from plants and other items ingested as food to those used for medicinal purpose, requiring an interdisciplinary approach integrating the compilation of herbaria, analyses of plant’s pharmacological properties, and their effect on the health status of individual bonobos.




2013, Ludwig Maximilian University of Munich, Germany, Habilitation
1995, Ludwig Maximilian University of Munich, Germany, Dr. rer.nat.


Journal article

Lee SM, Murray CM, Lonsdorf EV, Fruth B, Stanton MA, Nichols J, Hohmann G. 2020. Wild bonobo and chimpanzee females exhibit broadly similar patterns of behavioral maturation but some evidence for divergence American Journal of Physical Anthropology, 171 :100-109 DOI Author Url

Moscovice LR, Surbeck M, Fruth B, Hohmann G, Jaeggi AV, Deschner T. 2019. The cooperative sex: Sexual interactions among female bonobos are linked to increases in oxytocin, proximity and coalitions Hormones and Behaviour, 116 DOI Author Url Publisher Url Public Url

Hohmann G, Ortmann S, Remer T, Fruth B. 2019. Fishing for iodine: What aquatic foraging by bonobos tells us about human evolution BMC Zoology, DOI Author Url Publisher Url Public Url

Murthy S, O'Brien K, Agbor A, Angedakin S, Arandjelovic M, Ayimisin EA, Bailey E, Bergl RA, Brazzola G, Dieguez P, Eno-Nku M, Eshuis H, Fruth B, Gillespie TR, Ginath Y, Gray M, Herbinger I, Jones S, Kehoe L, Kuhl H, Kujirakwinja D, Lee K, Madinda NF, Mitamba G, Muhindo E, Nishuli R, Ormsby LJ, Petrzelkova KJ, Plumptre AJ, Robbins MM, Sommer V, Ter Heegde M, Todd A, Tokunda R, Wessling E, Jarvis MA, Leendertz FH, Ehlers B, Calvignac-Spencer S. 2019. Cytomegalovirus distribution and evolution in hominines Virus Evolution, 5 DOI Author Url Publisher Url Public Url

Surbeck M, Boesch C, Crockford C, Thompson ME, Furuichi T, Fruth B, Hohmann G, Ishizuka S, Machanda Z, Muller MN, Pusey A, Sakamaki T, Tokuyama N, Walker K, Wrangham R, Wroblewski E, Zuberbuhler K, Vigilant L, Langergraber K. 2019. Males with a mother living in their group have higher paternity success in bonobos but not chimpanzees CURRENT BIOLOGY, 29 :R354-R355 DOI Author Url

Kalan AK, Hohmann G, Arandjelovic M, Boesch C, McCarthy MS, Agbor A, Angedakin S, Bailey E, Belongelwa CW, Bessone M, Bocksberger G, Coxe SJ, Deschner T, Despres-Einspenner ML, Dieguez P, Fruth B, Herbinger I, Granjon AC, Head J, Kablan YA, Langergraber KE, Lokasola AL, Maretti G, Marrocoli S, Mbende M, Moustgaard J, N'Goran PK, Robbins MM, van Schijndel J, Sommer V, Surbeck M, Tagg N, Willie J, Wittig RM, Kuhl HS. 2019. Novelty Response of Wild African Apes to Camera Traps Current Biology, DOI Author Url Publisher Url Public Url

Fruth B, Tagg N, Stewart F. 2018. Sleep and nesting behavior in primates: A review American Journal of Physical Anthropology, 166 :499-509 DOI Author Url Publisher Url Public Url

Fruth B, Hohmann G. 2018. Food Sharing across Borders Human Nature: an interdisciplinary biosocial perspective, 29 :91-103 DOI Author Url Publisher Url Public Url

Bondjengo N, Kitengie G, Musibono D, Lubini C, Hohmann G, Fruth B. 2017. Presence of Alkaloids and Cyanogenic Glycosides in Fruits Consumed by Sympatric Bonobos and the Nkundo People at LuiKotale/Salonga National Park, Democratic Republic of Congo and Its Relationship to Food Choice African Primates, 12 :9-22 Publisher Url Public Url

Beaune D, Hohmann G, Serckx A, Sakamaki T, Narat V, Fruth B. 2017. How bonobo communities deal with tannin rich fruits: Re-ingestion and other feeding processes BEHAVIOURAL PROCESSES, 142 :131-137 DOI Author Url

Surbeck M, Langergraber KE, Fruth B, Vigilant L, Hohmann G. 2017. Male reproductive skew is higher in bonobos than chimpanzees Current Biology, 27 :R640-R641 DOI Author Url Publisher Url Public Url

Surbeck M, Girard-Buttoz C, Boesch C, Crockford C, Fruth B, Hohmann G, Langergraber KE, Zuberbuhler K, Wittig RM, Mundry R. 2017. Sex-specific association patterns in bonobos and chimpanzees reflect species differences in cooperation Royal Society Open Science, 4 DOI Author Url Publisher Url Public Url

Mossoun A, Calvignac-Spencer S, Anoh AE, Pauly MS, Driscoll DA, Michel AO, Nazaire LG, Pfister S, Sabwe P, Thiesen U, Vogler BR, Wiersma L, Muyembe-Tamfum J-J, Karhemere S, Akoua-Koffi C, Couacy-Hymann E, Fruth B, Wittig RM, Leendertz FH, Schubert G. 2017. Bushmeat Hunting and Zoonotic Transmission of Simian T-Lymphotropic Virus 1 in Tropical West and Central Africa JOURNAL OF VIROLOGY, 91 DOI Author Url

Muganza DM, Fruth B, Nzunzu JL, Tuenter E, Foubert K, Cos P, Maes L, Kanyanga RC, Exarchou V, Apers S, Pieters L. 2016. In vitro antiprotozoal activity and cytotoxicity of extracts and isolated constituents from Greenwayodendron suaveolens JOURNAL OF ETHNOPHARMACOLOGY, 193 :510-516 DOI Author Url

Madinda NF, Ehlers B, Wertheim JO, Akoua-Koffi C, Bergl RA, Boesch C, Akonkwa DBM, Eckardt W, Fruth B, Gillespie TR, Gray M, Hohmann G, Karhemere S, Kujirakwinja D, Langergraber K, Muyembe J-J, Nishuli R, Pauly M, Petrzelkova KJ, Robbins MM, Todd A, Schubert G, Stoinski TS, Wittig RM, Zuberbuhler K, Peeters M, Leendertz FH, Calvignac-Spencer S. 2016. Assessing Host-Virus Codivergence for Close Relatives of Merkel Cell Polyomavirus Infecting African Great Apes Journal of Virology, 90 :8531-8541 DOI Author Url Publisher Url Public Url

Oelze VM, Douglas PH, Stephens CR, Surbeck M, Behringer V, Richards MP, Fruth B, Hohmann G. 2016. The Steady State Great Ape? Long Term Isotopic Records Reveal the Effects of Season, Social Rank and Reproductive Status on Bonobo Feeding Behavior PLoS ONE, 11 DOI Author Url Publisher Url Public Url

Fruth BI, Hickey JR, André C, Furuichi T, Hart J, Hart T, Kuehl H, Maisels F, Nackoney J, Reinartz G, Sop T, Thompson J, Williamson EA. 2016. Pan paniscus The IUCN Red List of Threatened Species,

Goldstone LG, Sommer V, Nurmi N, Stephens C, Fruth B. 2016. Food begging and sharing in wild bonobos (Pan paniscus): assessing relationship quality? PRIMATES, 57 :367-376 DOI Author Url

Froehlich M, Kuchenbuch P, Mueller G, Fruth B, Furuichi T, Wittig RM, Pika S. 2016. Unpeeling the layers of language: Bonobos and chimpanzees engage in cooperative turn-taking sequences Scientific Reports, 6 DOI Author Url Publisher Url Public Url

Muganza DM, Fruth BI, Lami JN, Cos R, Kanyanga RC, Maes L, Pieters L. 2015. In vitro antiprotozoal activity and cytotoxicity of extracts and fractions from the leaves, root bark and stem bark of Isolona hexaloba JOURNAL OF ETHNOPHARMACOLOGY, 174 :187-194 DOI Author Url

Hoppe E, Pauly M, Gillespie TR, Akoua-Koffi C, Hohmann G, Fruth B, Karhemere S, Madinda NF, Mugisha L, Muyembe J-J, Todd A, Petrzelkova KJ, Gray M, Robbins M, Bergl RA, Wittig RM, Zuberbuehler K, Boesch C, Schubert G, Leendertz FH, Ehlers B, Calvignac-Spencer S. 2015. Multiple Cross-Species Transmission Events of Human Adenoviruses (HAdV) during Hominine Evolution MOLECULAR BIOLOGY AND EVOLUTION, 32 :2072-2084 DOI Author Url

Beaune D, Bretagnolle F, Bollache L, Hohmann G, Fruth B. 2015. Can fruiting plants control animal behaviour and seed dispersal distance? BEHAVIOUR, 152 :359-374 DOI Author Url

Wilson ML, Boesch C, Fruth B, Furuichi T, Gilby IC, Hashimoto C, Hobaiter CL, Hohmann G, Itoh N, Koops K, Lloyd JN, Matsuzawa T, Mitani JC, Mjungu DC, Morgan D, Muller MN, Mundry R, Nakamura M, Pruetz J, Pusey AE, Riedel J, Sanz C, Schel AM, Simmons N, Waller M, Watts DP, White F, Wittig RM, Zuberbuehler K, Wrangham RW. 2014. Lethal aggression in Pan is better explained by adaptive strategies than human impacts NATURE, 513 :414-+ DOI Author Url

Fruth B, Ikombe NB, Matshimba GK, Metzger S, Muganza DM, Mundry R, Fowler A. 2014. New Evidence for Self-Medication in Bonobos: Manniophyton fulvum Leaf- and Stemstrip-Swallowing From LuiKotale, Salonga National Park, DR Congo AMERICAN JOURNAL OF PRIMATOLOGY, 76 :146-158 DOI Author Url

Beaune D, Bretagnolle F, Bollache L, Bourson C, Hohmann G, Fruth B. 2013. Ecological services performed by the bonobo (Pan paniscus): seed dispersal effectiveness in tropical forest JOURNAL OF TROPICAL ECOLOGY, 29 :367-380 DOI Author Url

Beaune D, Bollache L, Muganza MD, Bretagnolle F, Hohmann G, Fruth B. 2013. Artificial Germination Activation of Dialium corbisieri by Imitation of Ecological Process Journal of Sustainable Forestry, 32 :565-575 DOI

Beaune D, Fruth B, Bollache L, Hohmann G, Bretagnolle F. 2013. Doom of the elephant-dependent trees in a Congo tropical forest FOREST ECOLOGY AND MANAGEMENT, 295 :109-117 DOI Author Url

Beaune D, Bretagnolle F, Bollache L, Hohmann G, Surbeck M, Bourson C, Fruth B. 2013. The Bonobo-Dialium Positive Interactions: Seed Dispersal Mutualism AMERICAN JOURNAL OF PRIMATOLOGY, 75 :394-403 DOI Author Url

Beaune D, Bretagnolle F, Bollache L, Hohmann G, Surbeck M, Fruth B. 2013. Seed dispersal strategies and the threat of defaunation in a Congo forest BIODIVERSITY AND CONSERVATION, 22 :225-238 DOI Author Url

Fruth B, Hohmann G. 2013. The Nature of the Bonobo Society: A Case Study on the Learning Process of Primatologists FOLIA PRIMATOLOGICA, 84 :274-274 Author Url

Beaune D, Bollache L, Fruth B, Bretagnolle F. 2012. Bush pig (Potamochoerus porcus) seed predation of bush mango (Irvingia gabonensis) and other plant species in Democratic Republic of Congo AFRICAN JOURNAL OF ECOLOGY, 50 :509-512 DOI Author Url

Laurance WF, Useche DC, Rendeiro J, Kalka M, Bradshaw CJA, Sloan SP, Laurance SG, Campbell M, Abernethy K, Alvarez P, Arroyo-Rodriguez V, Ashton P, Benitez-Malvido J, Blom A, Bobo KS, Cannon CH, Cao M, Carroll R, Chapman C, Coates R, Cords M, Danielsen F, De Dijn B, Dinerstein E, Donnelly MA, Edwards D, Edwards F, Farwig N, Fashing P, Forget P-M, Foster M, Gale G, Harris D, Harrison R, Hart J, Karpanty S, Kress WJ, Krishnaswamy J, Logsdon W, Lovett J, Magnusson W, Maisels F, Marshall AR, McClearn D, Mudappa D, Nielsen MR, Pearson R, Pitman N, van der Ploeg J, Plumptre A, Poulsen J, Quesada M, Rainey H, Robinson D, Roetgers C, Rovero F, Scatena F, Schulze C, Sheil D, Struhsaker T, Terborgh J, Thomas D, Timm R, Urbina-Cardona JN, Vasudevan K, Wright SJ, Arias-G JC, Arroyo L, Ashton M, Auzel P, Babaasa D, Babweteera F, Baker P, Banki O, Bass M, Bila-Isia I, Blake S, Brockelman W, Brokaw N, Bruehl CA, Bunyavejchewin S, Chao J-T, Chave J, Chellam R, Clark CJ, Clavijo J, Congdon R, Corlett R, Dattaraja HS, Dave C, Davies G, Beisiegel BDM, da Silva RDN, Di Fiore A, Diesmos A, Dirzo R, Doran-Sheehy D, Eaton M, Emmons L, Estrada A, Ewango C, Fedigan L, Feer F, Fruth B, Willis JG, Goodale U, Goodman S, Guix JC, Guthiga P, Haber W, Hamer K, Herbinger I, Hill J, Huang Z, Sun IF, Ickes K, Itoh A, Ivanauskas N, Jackes B, Janovec J, Janzen D, Jiangming M, Jin C, Jones T, Justiniano H, Kalko E, Kasangaki A, Killeen T, King H-B, Klop E, Knott C, Kone I, Kudavidanage E, Ribeiro JLDS, Lattke J, Laval R, Lawton R, Leal M, Leighton M, Lentino M, Leonel C, Lindsell J, Ling-Ling L, Linsenmair KE, Losos E, Lugo A, Lwanga J, Mack AL, Martins M, McGraw WS, McNab R, Montag L, Thompson JM, Nabe-Nielsen J, Nakagawa M, Nepal S, Norconk M, Novotny V, O'Donnell S, Opiang M, Ouboter P, Parker K, Parthasarathy N, Pisciotta K, Prawiradilaga D, Pringle C, Rajathurai S, Reichard U, Reinartz G, Renton K, Reynolds G, Reynolds V, Riley E, Roedel M-O, Rothman J, Round P, Sakai S, Sanaiotti T, Savini T, Schaab G, Seidensticker J, Siaka A, Silman MR, Smith TB, de Almeida SS, Sodhi N, Stanford C, Stewart K, Stokes E, Stoner KE, Sukumar R, Surbeck M, Tobler M, Tscharntke T, Turkalo A, Umapathy G, van Weerd M, Rivera JV, Venkataraman M, Venn L, Verea C, de Castilho CV, Waltert M, Wang B, Watts D, Weber W, West P, Whitacre D, Whitney K, Wilkie D, Williams S, Wright DD, Wright P, Xiankai L, Yonzon P, Zamzani F. 2012. Averting biodiversity collapse in tropical forest protected areas NATURE, 489 :290-+ DOI Author Url

Beaune D, Bollache L, Bretagnolle F, Fruth B. 2012. Dung beetles are critical in preventing post-dispersal seed removal by rodents in Congo rain forest JOURNAL OF TROPICAL ECOLOGY, 28 :507-510 DOI Author Url

Beaune D, Bollache L, Fruth B, Hohmann G, Bretagnolle F. 2012. Density-dependent effect affecting elephant seed-dispersed tree recruitment (Irvingia gabonensis) in Congo Forest PACHYDERM, :97-100 Author Url

Muganza DM, Fruth BI, Lami JN, Mesia GK, Kambu OK, Tona GL, Kanyanga RC, Cos P, Maes L, Apers S, Pieters L. 2012. In vitro antiprotozoal and cytotoxic activity of 33 ethonopharmacologically selected medicinal plants from Democratic Republic of Congo JOURNAL OF ETHNOPHARMACOLOGY, 141 :301-308 DOI Author Url

Fruth BI. 2012. La Convention sur la Diversité Biologique en République Démocratique du Congo : Mise en oeuvre du projet Ethnopharmacologia, 48 :27-37

Oelze VM, Fuller BT, Richards MP, Fruth B, Surbeck M, Hublin J-J, Hohmann G. 2011. Exploring the contribution and significance of animal protein in the diet of bonobos by stable isotope ratio analysis of hair PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA, 108 :9792-9797 DOI Author Url

Fruth BI. 2011. The CBD in the Democratic Republic of Congo (DRC): the project “The Cuvette Centrale as a reservoir of medicinal plants” in the process of implementation. Curare -Berlin-, 34 :51-62

Fruth BI. 2011. Care for health and body: an ethnobotanical approach to Nkundo plant use (Cuvette Centrale, DRC) with focus on the significance of indigenous knowledge for the human skin. Curare -Berlin-, 34 :261-281

Hohmann G, Fruth B. 2008. New records on prey capture and meat eating by bonobos at Lui Kotale, Salonga National Park, Democratic Republic of Congo FOLIA PRIMATOLOGICA, 79 :103-110 DOI Author Url

McGrew WC, Marchant LF, Beuerlein MM, Vrancken D, Fruth B, Hohmann G. 2007. Prospects for bonobo insectivory: Lui kotal, democratic republic of congo INTERNATIONAL JOURNAL OF PRIMATOLOGY, 28 :1237-1252 DOI Author Url

D'Amour DE, Hohmann G, Fruth B. 2006. Evidence of leopard predation on bonobos (Pan paniscus) FOLIA PRIMATOLOGICA, 77 :212-217 DOI Author Url

Fruth BI. 2006. Grooming hand clasp by bonobos of Lui Kotal, Democratic Republic of Congo Pan Africa News, 13 :6-8

Boubli JP, Eriksson J, Wich S, Hohmann G, Fruth B. 2004. Mesoscale transect sampling of trees in the Lomako-Yekokora interfluvium, Democratic Republic of the Congo BIODIVERSITY AND CONSERVATION, 13 :2399-2417 DOI Author Url

Fruth B, Hohmann G. 2004. Determinants of spatial associations in a co-dominant society: Variation of fission-fusion patterns in bonobos FOLIA PRIMATOLOGICA, 75 :139-140 Author Url

Hohmann G, Fruth B. 2003. Intra- and inter-sexual aggression by bonobos in the context of mating BEHAVIOUR, 140 :1389-1413 DOI Author Url

Hohmann G, Fruth B. 2003. Culture in Bonobos? Between-species and within-species variation in behavior CURRENT ANTHROPOLOGY, 44 :563-571 DOI Author Url

Fruth BI. 2003. Lui Kotal - A new site for field research on bonobos in the Salonga National Park Pan Africa News, 10 :25-27

Hohmann G, Fruth B. 2000. Use and function of genital contacts among female bonobos ANIMAL BEHAVIOUR, 60 :107-120 DOI Author Url

Hohmann G, Gerloff U, Tautz D, Fruth B. 1999. Social bonds and genetic ties: Kinship association and affiliation in a community of bonobos (Pan paniscus) BEHAVIOUR, 136 :1219-1235 DOI Author Url

Gerloff U, Hartung B, Fruth B, Hohmann G, Tautz D. 1999. Intracommunity relationships, dispersal pattern and paternity success in a wild living community of Bonobos (Pan paniscus) determined from DNA analysis of faecal samples PROCEEDINGS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES, 266 :1189-1195 DOI Author Url

Gagneux P, Wills C, Gerloff U, Tautz D, Morin PA, Boesch C, Fruth B, Hohmann G, Ryder OA, Woodruff DS. 1999. Mitochondrial sequences show diverse evolutionary histories of African hominoids PROCEEDINGS OF THE NATIONAL ACADEMY OF SCIENCES OF THE UNITED STATES OF AMERICA, 96 :5077-5082 DOI Author Url

Fruth B, McGrew WC. 1998. Resting and nesting in primates: Behavioral ecology of inactivity AMERICAN JOURNAL OF PRIMATOLOGY, 46 :3-5 DOI Author Url

Marchesi P, Marchesi N, Fruth B, Boesch C. 1995. Census and distribution of chimpanzees in Côte D'Ivoire Primates, 36 :591-607 DOI


Fruth BI. 1995. Nestbuilding behavior in captive bonobos (Pan paniscus) Primate report, 42 :43-45



Fruth BI. 1994. Is nut cracking in wild chimpanzees a cultural behaviour? Journal of Human Evolution, 26 :325-338 DOI

Fruth BI, Hohmann G. 1993. Ecological and behavioural aspects of nest building in wild bonobos (Pan paniscus). Ethology, 94 :113-126 DOI


Payne C, Mato B, Fruth B. Entomophagy in the area surrounding LuiKotale, Salonga National Park, Democratic Republic of the Congo African Study Monographs,, 1 :1-12 DOI Public Url

Conference publication

Moscovice LR, Surbeck M, Fruth B, Hohmann G, Deschner T. 2018. The Sex that Binds: Genito-genital rubbing is associated with increases in urinary oxytocin among wild female bonobos AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, 87th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA) 165 :182-183 Author Url

Jaeggi AV, Moscovice LR, Goldstone LG, Hohmann G, Fruth B. 2017. Reciprocity can Explain Grooming, but not other Forms of Cooperation, among Female Bonobos at LuiKotale, DRC AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, 86th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA) 162 :229-229 Author Url

HOHMANN G, FRUTH B. 1995. Loud calls in great apes: Sex differences and social correlates Zimmermann E, Newman JD, Jurgens U. CURRENT TOPICS IN PRIMATE VOCAL COMMUNICATION, XIV Congress of the International-Primatological-Society :161-184 Author Url

Fruth BI, Muganza M. Traditional use of wild rainforest plants by the Nkundo, Democratic Republic of Congo (DRC) Proceedings of the IVth International Congress of Ethnobotany, IVth International Congress of Ethnobotany (ICEB, 2005). :471-473


Fruth BI. 2016. Great Ape Nest-Building Fuentes A. The International Encyclopedia of Primatology. 1-3 9781119179313 DOI

Hohmann G, Fruth B. 2011. Is blood thicker than water? Among African Apes: Stories and Photos from the Field :61-76 9780520267107

Fruth BI, Mato B, Muganza M. 2010. Nkundo plant use (Cuvette Centrale, DRC) with a focus on indigenous knowledge and the application of aphrodisiacs Building Bridges between Anthropology, Medicine and Human Ethology :211-226 Bochum: The University Press 9783899663488

Mohneke M, Fruth BI. 2007. Bonobo (Pan paniscus) density estimation in the SW-Salonga National Park, Democratic Republic of Congo: common methodology revisited. The Bonobos Behavior, Ecology, and Conservation :151-166 Springer Science & Business Media 9780387747873 DOI Author Url

Fruth BI, Hohmann G. 2006. Social Grease for females? Same sex genital contacts in wild bonobos Homosexual Behaviour in Animals An Evolutionary Perspective Cambridge University Press 9780521864466

Fruth BI, Hohmann G. 2002. How bonobos handle hunts and harvests: why share food? Behavioural Diversity in Chimpanzees and Bonobos :231-243 Cambridge University Press 9780521006132

Fruth BI. 2002. Dynamics in social organization of bonobos (Pan paniscus) Behavioural Diversity in Chimpanzees and Bonobos :138-150 Cambridge University Press 9780521006132

Fruth BI. 1999. The Pan species The Nonhuman Primates McGraw-Hill Humanities Social 978-1559349741

Fruth BI, Hohmann G. 1996. Comparative analyses of nest building behavior in bonobos (Pan paniscus) and chimpanzees (Pan troglodytes Chimpanzee Cultures :109-128 Harvard University Press 9780674116634

Fruth BI, Hohmann G. 1996. Nest building behavior in the great apes: the great leap forward? Great Ape Societies :225-240 Cambridge University Press 9780521555364

Fruth BI. Das Kongobecken und seine Regenwälder in Vergangenheit, Gegenwart und Zukunft. Feest C, Kron C. Regenwald :110-119 Konrad Theiss. Wemding 978-3-8062-2799-4

Fruth BI. Primaten: Verwandtschaft aus dem Regenwald Feest C, Kron C. Regenwald :130-135 Konrad Theiss. Wemding 978-3-8062-2799-4

Williamson EA, Maisels FG, Groves CP, Fruth BI, Humle T, Morton B, Richardson MC, Russon AE, Singleton I. Family Hominidae (Great Apes) Handbook of the Mammals of the World Primates :853-854 Lynx Edicions 9788496553897

Liebs V, Bikandu B, Lassa L, Lukoki L, Schareika N, Fruth BI. Signifying competence: urban herbalists seeking affirmation in Kinshasa Living the City in Africa Processes of Invention and Intervention :213-233 LIT Verlag Münster 9783643801524

Fruth BI, Hohmann G. Food sharing and status in unprovisioned bonobos Food and the Status Quest An Interdisciplinary Perspective :47-67 Berghahn Books 9781571811233

Fruth B, Hohmann G. Loud calls in great apes: Sex differences and social correlates Current Topics in Primate Vocal Communication :161-184 Springer Science & Business Media 9781475799309 DOI

Engagement & Impact

Research Grants Awarded:

LJMU SEED CORN funding, Development of wild bonobos., Grant value (£): 10000, Duration of research project: long-term. 2019

Leakey Foundation, Emergency funding to support/maintain the site of LuiKotale, Grant value (£): US$ 25.000, Duration of research project: 1 year. 2018

CRC / KMDA, LuiKotale Bonobo Project (LKBP), Grant value (£): € 475.000, Duration of research project: 5 years. 2017

KfW / WWF, Proposition d’un système de suivi écologique comme outil de gestion de la Salonga Sud, RDC: Inventaire de la biodiversité et développement d’une stratégie pour améliorer la protection de la faune et flore menacées, Grant value (£): € 587000, Duration of research project: 18 months. 2016

CRC/RZSA, LuiKotale Bonobo Project (LKBP), Grant value (£): €95.000 (per annum), Duration of research project: 2 years. 2014

ARCUS Foundation, Viability assessment of Bonobo Ecotourism, Grant value (£): $156,356, Duration of research project: 2 years. 2013

USFWS, Improving in-situ conservation of bonobos (Pan paniscus) by environmental education and capacity building in schools and villages West of Salonga National Park, DRC, Grant value (£): US$ 55.585, Duration of research project: 2 years. 2011

USFWS, Improving in-situ conservation of bonobos (Pan paniscus) by environmental education and capacity building in schools and villages West of Salonga National Park, DRC, Grant value (£): $55,585. 2011

German Embassy Kinshasa (DBO-KIN), Establishment of identification plates of traditional medicinal plants for medicinal garden, Grant value (£): $1,800, Duration of research project: 1 year. 2010

German Research Council (DFG), Capacity Building for the Detection and Prevention of Neglected Zoonotic Diseases in Tropical Africa, PI Dr. Fabian Leendertz, Grant value (£): € 398,845, Duration of research project: 3 years. 2010

Federal Ministry for Education & Research (BMBF), The Cuvette Centrale as reservoir of medicinal plants, Grant value (£): €590,000, Duration of research project: 3 years. 2007

Other invited event:

Medical Anthropology seminar series, School of Anthropology & Museum Ethnography, Oxford University, UK, Talk: “Self-medication in Humans and Bonobos: Lessons learned from Congo's Wild Pharmacy”. 2018

Workshop “Development of Biomonitoring Strategy” ICCN/WWF-DRC, Kinshasa, DRC, Talk: "Résultats d’Inventaire du Park National de la Salonga Sud: Faune, Flore et Analyse Satellitaire". 2018

Pint of Science, Munich; Germany, Talk: “The bonobo: an ape that shapes our future”. 2018

38th Workshop of the Association of Zoo Veterinarians, Stuttgart, Germany, Keynote talk: “Eigenmedikation bei Menschenaffen: Eine Suche nach dem Arzt im Tier”. 2018

Studium Generale, Johannes Gutenberg University of Mainz, Lecture: “Der kongolesische Regenwald als Ressource für Mensch und Tier”. 2018

Symposium "Challenges in Modern Biology", Max Planck Institute for Evolutionary Biology, Plön, Germany, Talk: "Bonobos are different: Understanding female power". 2017

Seminar, Royal Zoological Society of Antwerp, Belgium, Talk: “Bonobo research: Visions towards a ‘One Plan Approach’". 2017

Bonobos and the Congo’s Wild Pharmacy: Hints towards the Evolution of Self-Medication?, Muséum national d'Histoire naturelle (MNHN), Paris, Invited talk. 2014

BONOBOS, unsere nächsten Verwandten, NaMu Science after Work, München, Invited talk. 2014

Bonobos and the Congo's Wild Pharmacy: Hints towards the evolution of self-medication?, Belgian Group for Primatology (BGP), Brussels, Meeting of Primatology, Invited talk. 2014

Evolution of Man: "Medicinal plant use in the Cuvette Centrale of the DR Congo: Ethnobotanical Investigations in Rural and Urban Environments, Center for Advanced Studies (CAS), LMU-Munich, Invited talk. 2014

Das Kongobecken, Urwaldapotheke für Mensch und Tier: Was können wir von Bonobos lernen?, Senior-academy, LMU-Munich, Invited talk. 2014

The Evolution of Self-medication: Humans, bonobos, and the Congo's wild pharmacy, BioAnthro-Seminar talks, Dept. of Anthropology, UCL London, Invited talk. 2014

Specialists shaped by nature: what HEFA can tell us about our past, present and future, Senckenberg Research Institute, Frankfurt, Workshop: Human Nature & Culture: Unity, Diversity and Change in the Human Niches. 2014

Freilandforschung und Naturschutz im Kongo (DRC): Bonobos" (Research & conservation in Congo, DRC), Zoological Garden, Cologne, Seminar-Series. 2014

The nature of the bonobo society: a case study on the learning process of primatologists, European Federation for Primatology meeting, Antwerp, Belgium, Keynote. 2013

Bonobos im Kongo, Zoo-association Wuppertal, Germany, Seminar-Series. 2013

Wie Menschen Affen sehen (How humans see apes), Neanderthal Museum, Mettmann, Germany, Symposium. 2012

Die Taschen der Tiere - Einblick in die Evolution des Tragens (Animal bags - insight into the evolution of carrying), Deutsches Hygiene-Museum, Dresden, Germany, Symposium: Homo portans: Tragen, die Faszination des Selbstverständlichen (Homo portans: Carrying, fascination of the obvious). 2011

External committees:

Thesis committee, Georg-August University Göttingen, Germany, Examinor PhD-Thesis "Medicinal Plant Use by Herbalists and Lay-people in an Urban Context: Problems and Options in Kinshasa (DRC)" V. Liebs. 2017

Thesis Committee, Antwerp University, Belgium, Examiner; PhD-Thesis “Ethnobotanical, Phytochemical and Biological Investigations on Medicinal Plants Used by the Nkundo (Bandundu, DR Congo)” C.D.M.Muganza. 2017

Thesis committee, Muséum national d'Histoire naturelle, Examiner. 2014

Thesis Committee: Behavioral Biology Unit, University of Liege, Belgium, Referee and Examiner. 2014

HDR Thesis committee; Les chimpanzés de Kibale (Ouganda): Santé, usage de substances naturelles et interactions avec les humains; Habilitation à Diriger des Recherches, Université de Bourgogne, France, Referee and Examiner. 2012

Thesis committee: The Evolution of Shelter: Ecology and Evolution of Chimpanzee Nest Building, University of Cambridge, UK, Referee and Examiner.

Membership of professional bodies:

Executive Director, Bonobo Alive, 2011

Member of Executive Committee, Great Ape Section (SGA) of the IUCN/SSC Primate Specialist Group, 2009

Media Coverage:, The LuiKotale Bonobo Project; Salonga National Park, DRC.,,, Publication: Fruth B. & G. Hohmann (2018) Food Sharing across borders: First observation of inter-community meat sharing by bonobos at LuiKotale, DRC. Human Nature, 29(2), 91-103.., WWF BIOMONITORING IN SALONGA NATIONAL PARK, DR CONGO.