Prof Peter Falkingham

School of Biological and Environmental Sciences

Reader in Vertebrate Biology

Please visit my personal website for my blog and a much clearer overview of my research and publications: https://peterfalkingham.com

My primary research focus is on dinosaur locomotion and footprint formation. Specifically, how the foot and sediment interact, and how we can subsequently reverse-engineer their formation in order to constrain and understand the limb motion of these extinct animals.

By understanding the motion of substrate around a dynamically moving foot, it is possible to use tracks to constrain possible motions of the distal (and therefore proximal) limb. My work incorporates digitisation of fossil tracks and computer simulation as well as using data from extant taxa using XROMM techniques to study the sub-surface motion of the foot during locomotion.

In addition to my work on footprints, I’m also involved in other research areas including dinosaur biomechanics and taxonomy, development of digitsation techniques and applications, cambrian echinoderm hydrodynamics, and materials science, collaborating with scientists from the UK, USA, across Europe, Australia, and South America.

Languages

English

Degrees

2010, University of Manchester, United Kingdom, PhD Palaeontology - Computer Simulation of Dinosaur Tracks
2004, University of Bristol, United Kingdom, Computer Science MSc
2003, University of Bristol, UK, Biology and Geology BSc (Hons)

Academic appointments

Reader in Vertebrate Biology, School of Biological and Environmental Sciences, Liverpool John Moores University, 2020 - present
Lecturer/Senior Lecturer Vertebrate Biology, Natural Sciences and Psychology, Liverpool John Moores University, 2015 - 2020
Marie Curie Research Fellow, Comparative Biomedical Sciences, Royal Veterinary College, 2012 - 2015

Postgraduate training

Marie Curie International Outgoing Fellow, UK/USA, Royal Veterinary College/Brown University, 2012 - 2015

Highlighted publications

Hatala KG, Gatesy SM, Falkingham PL. 2023. Arched footprints preserve the motions of fossil hominin feet Nature Ecology and Evolution, DOI Author Url Publisher Url Public Url

Lallensack JN, Romilio A, Falkingham PL. 2022. A machine learning approach for the discrimination of theropod and ornithischian dinosaur tracks Journal of the Royal Society Interface, 19 DOI Author Url Publisher Url Public Url

Turner ML, Falkingham PL, Gatesy SM. 2022. What is Stance Phase On Deformable Substrates? Integrative and Comparative Biology, DOI Publisher Url Public Url

Lallensack JN, Falkingham PL. 2022. A new method to calculate limb phase from trackways reveals gaits of sauropod dinosaurs Current Biology, DOI Author Url Publisher Url Public Url

Falkingham P, Turner M, Gatesy S. 2020. Constructing and testing hypotheses of dinosaur foot motions from fossil tracks using digitization and simulation Palaeontology, DOI Author Url Publisher Url Public Url

Tuner M, Falkingham PL, Gatesy S. 2020. It's in the loop: shared sub-surface foot kinematics in birds and other dinosaurs shed light on a new dimension of fossil track diversity Biology Letters, 16 DOI Author Url Publisher Url Public Url

Crandell K, Howe R, Falkingham P. 2019. Repeated evolution of drag reduction at the air-water interface in diving kingfishers Journal of the Royal Society Interface, 16 DOI Author Url Publisher Url Public Url

Gatesy SM, Falkingham PL. 2017. Neither bones nor feet: track morphological variation and “preservation quality” Journal of Vertebrate Paleontology, DOI Author Url Publisher Url Public Url

Rahman I, Zamora S, Falkingham PL, Phillips JC. 2015. Cambrian cinctan echinoderms shed light on feeding in the ancestral deuterostome Proceedings of the Royal Society B: Biological Sciences, 282 DOI Author Url Publisher Url Public Url

Falkingham PL, Gatesy SM. 2014. The birth of a dinosaur footprint: Subsurface 3D motion reconstruction and discrete element simulation reveal track ontogeny Proceedings of the National Academy of Sciences of the United States of America, 111 :18279-18284 DOI Author Url Publisher Url Public Url

Falkingham PL. 2014. Interpreting ecology and behaviour from the vertebrate fossil track record JOURNAL OF ZOOLOGY, 292 :222-228 DOI Author Url Publisher Url

Journal article

Hatala KG, Gatesy SM, Manafzadeh AR, Lusardi E, Falkingham PL. 2024. Technical note: A volumetric method for measuring the longitudinal arch of human tracks and feet American Journal of Biological Anthropology, DOI Author Url Publisher Url Public Url

Oussou A, Falkingham PL, Butler RJ, Boumir K, Ouarhache D, Ech-charay K, Charrière A, Maidment SCR. 2023. New Middle to ?Late Jurassic dinosaur tracksites in the Central High Atlas Mountains, Morocco Royal Society Open Science, 10 DOI Author Url Publisher Url Public Url

Vincelette A, Renders E, Scott K, Falkingham P, Janis C. 2023. Hipparion tracks and horses' toes: the evolution of the equid single hoof Royal Society Open Science, 10 DOI Author Url Publisher Url Public Url

Flannery-Sutherland JT, Kogan I, Trubin YS, Falkingham PL, Winkler A, De Sousa DD, Krylov KD, Pokhaznikova AA, Derbisheva M, Kapitany T, Dudashvili A. 2023. Dinosaur trackways from the Upper Cretaceous Nichkesai Formation near Mayluu Suu City, Southern Tien Shan Mountains, north-western Kyrgyzstan Royal Society Open Science, 10 DOI Publisher Url Public Url

Hatala KG, Gatesy SM, Falkingham PL. 2023. Arched footprints preserve the motions of fossil hominin feet Nature Ecology and Evolution, DOI Author Url Publisher Url Public Url

Grant B, Charles J, Geraghty B, Gardiner J, D'Août K, Falkingham PL, Bates KT. 2022. Why does the metabolic cost of walking increase on compliant substrates? Journal of The Royal Society Interface, 19 DOI Author Url Publisher Url Public Url

Lallensack JN, Romilio A, Falkingham PL. 2022. A machine learning approach for the discrimination of theropod and ornithischian dinosaur tracks Journal of the Royal Society Interface, 19 DOI Author Url Publisher Url Public Url

Lallensack JN, Owais A, Falkingham PL, Breithaupt BH, Sander PM. 2022. How to verify fossil tracks: the first record of dinosaurs from Palestine Historical Biology: an international journal of paleobiology, :1-11 DOI Publisher Url Public Url

Turner ML, Falkingham PL, Gatesy SM. 2022. What is Stance Phase On Deformable Substrates? Integrative and Comparative Biology, DOI Publisher Url Public Url

Lallensack JN, Falkingham PL. 2022. A new method to calculate limb phase from trackways reveals gaits of sauropod dinosaurs Current Biology, DOI Author Url Publisher Url Public Url

Falkingham PL, Maidment SCR, Lallensack JN, Martin JE, Suan G, Cherns L, Howells C, Barrett PM. 2021. Late Triassic dinosaur tracks from Penarth, south Wales Geological Magazine, :1-12 DOI Author Url Publisher Url Public Url

Lallensack JN, Farlow JO, Falkingham PL. 2021. A new solution to an old riddle: elongate dinosaur tracks explained as deep penetration of the foot, not plantigrade locomotion Palaeontology, DOI Author Url Publisher Url Public Url

Cieri RL, Turner ML, Carney RM, Falkingham PL, Kirk AM, Wang T, Jensen B, Novotny J, Tveite J, Gatesy SM, Laidlaw DH, Kaplan H, Moorman AFM, Howell M, Engel B, Cruz C, Smith A, Gerichs W, Lian Y, Schultz JT, Farmer CG. 2021. Virtual and augmented reality: New tools for visualizing, analyzing, and communicating complex morphology. Journal of Morphology, DOI Author Url Publisher Url Public Url

Farlow JO, Falkingham PL, Therrien F. 2021. Pedal proportions of small and large hadrosaurs and other potentially bipedal ornithischian dinosaurs CRETACEOUS RESEARCH, 127 DOI Author Url Publisher Url Public Url

Peri E, Falkingham PL, Collareta A, Bianucci G. 2021. Biting in the Miocene seas: estimation of the bite force of the macroraptorial sperm whale Zygophyseter varolai using finite element analysis Historical Biology: an international journal of paleobiology, DOI Author Url Publisher Url Public Url

Hatala KG, Gatesy SM, Falkingham PL. 2021. Integration of biplanar X-ray, three-dimensional animation and particle simulation reveals details of human ‘track ontogeny’ Interface Focus, 11 DOI Author Url Publisher Url Public Url

Mármol-Guijarro A, Nudds R, Folkow L, Sellers W, Falkingham P, Codd J. 2021. The Influence of snow properties on speed and gait choice in the Svalbard rock ptarmigan (Lagopus muta hyperborea) Integrative Organismal Biology, 3 DOI Author Url Publisher Url Public Url

Marek RD, Falkingham PL, Benson RBJ, Gardiner JD, Maddox TW, Bates KT. 2021. Evolutionary versatility of the avian neck Proceedings of the Royal Society B: Biological Sciences, 288 DOI Author Url Publisher Url Public Url

Tanaka I, Markó A, Hyodo M, Strickson C, Falkingham PL. 2021. A re-analysis of Chibanian Pleistocene tracks from VÉRTESSZŐLŐS, Hungary, employing photogrammetry and 3D analysis Annales Societatis Geologorum Poloniae, 91 :75-83 DOI Author Url Publisher Url Public Url

Falkingham PL, Rae R. 2021. 3D morphology of nematode encapsulation in snail shells, revealed by micro-CT imaging Scientific Reports, 11 DOI Author Url Publisher Url Public Url

Milàn J, Falkingham PL, Mueller-Töwe IJ. 2020. Small ornithopod dinosaur tracks and crocodilian remains from the Middle Jurassic Bagå Formation, Bornholm, Denmark: Important additions to the rare Middle Jurassic vertebrate faunas of Northern Europe Bulletin of the Geological Society of Denmark, :245-253 DOI Author Url Publisher Url Public Url

Gatesy SM, Falkingham PL. 2020. Hitchcock’s Leptodactyli, Penetrative Tracks, and Dinosaur Footprint Diversity Journal of Vertebrate Paleontology, DOI Author Url Publisher Url Public Url

Larkin NR, Duffin CJ, Dey S, Stukins S, Falkingham P. 2020. The first tetrapod track recorded from the Rhaetian in the British Isles Proceedings of the Geologists' Association, DOI Author Url Publisher Url Public Url

Falkingham P, Turner M, Gatesy S. 2020. Constructing and testing hypotheses of dinosaur foot motions from fossil tracks using digitization and simulation Palaeontology, DOI Author Url Publisher Url Public Url

Falkingham P, Gatesy S. 2020. Discussion: Defining the morphological quality of fossil footprints. Problems and principles of preservation in tetrapod ichnology with examples from the Palaeozoic to the present by Lorenzo Marchetti et al. Earth-Science Reviews, DOI Author Url Publisher Url

Tuner M, Falkingham PL, Gatesy S. 2020. It's in the loop: shared sub-surface foot kinematics in birds and other dinosaurs shed light on a new dimension of fossil track diversity Biology Letters, 16 DOI Author Url Publisher Url Public Url

Farlow JO, Richards RL, Johnson BD, Falkingham PL. 2020. A set-up for rapid screen-washing unconsolidated sediments for salvage-collecting large vertebrate fossils (pipe creek sinkhole, Late Neogene, Grant County, Indiana, USA) Proceedings Of The Indiana Academy Of Science, 129 :88-96

Otero A, Moreno AP, Falkingham PL, Cassini G, Ruella A, Militello M, Toledo N. 2020. Three-dimensional image surface acquisition in vertebrate paleontology: A review of principal techniques Publicación Electrónica de la Asociación Paleontológica Argentina, 20 DOI Publisher Url Public Url

Farlow JO, Bakker RT, Dattilo BF, Deschner EE, Falkingham PL, Harter C, Solis R, Temple D, Ward W. 2019. Thunder lizard handstands: Manus-only sauropod trackways from the Glen Rose Formation (Lower Cretaceous, Kendall County, Texas) Ichnos, DOI Author Url Publisher Url Public Url

Strickson EC, Hutchinson JR, Wilkinson DM, Falkingham PL. 2019. Can Skeletal Surface Area Predict in vivo Foot Surface Area? Journal of Anatomy, :72-84 DOI Author Url Publisher Url Public Url

Troelsen P, Wilkinson D, Seddighi M, Allanson D, Falkingham PL. 2019. Functional morphology and hydrodynamics of plesiosaur necks: Does size matter? Journal of Vertebrate Paleontology, 39 DOI Author Url Publisher Url Public Url

Crandell K, Howe R, Falkingham P. 2019. Repeated evolution of drag reduction at the air-water interface in diving kingfishers Journal of the Royal Society Interface, 16 DOI Author Url Publisher Url Public Url

Novotny J, Tveite J, Turner M, Gatesy S, Drury F, Falkingham PL, Laidlaw D. 2019. Developing Virtual Reality Visualizations for Unsteady Flow Analysis of Dinosaur Track Formation using Scientific Sketching IEEE Transactions on Visualization and Computer Graphics, 25 :2145-2154 DOI Author Url Publisher Url Public Url

Bates KT, Falkingham PL. 2018. The importance of muscle architecture in biomechanical reconstructions of extinct animals: a case study using Tyrannosaurus rex Journal of Anatomy, DOI Author Url Publisher Url Public Url

Falkingham PL, Bates KT, Avanzini M, Bennett M, Bordy E, Breithaupt BH, Castanera D, Citton P, Díaz-Martinez I, Farlow JO, Fiorillo AR, Gatesy SM, Getty P, Hatala KG, Hornung JJ, Hyatt JA, Lallensack JN, Martin AJ, Marty D, Matthews NA, Meyer CA, Milán J, Minter NJ, Razzolini NL, Romilio A, Salisbury SW, Sciscio L, Tanaka I, Wiseman ALA, Xing LD, Belvedere M. 2018. A standard protocol for documenting modern and fossil ichnological data Palaeontology, 61 :469-480 DOI Author Url Publisher Url Public Url

Gatesy SM, Falkingham PL. 2017. Neither bones nor feet: track morphological variation and “preservation quality” Journal of Vertebrate Paleontology, DOI Author Url Publisher Url Public Url

Lomax DR, Falkingham PL, Schweigert G, Jimenez AP. 2017. An 8.5 m long ammonite drag mark from the Upper Jurassic Solnhofen Lithographic Limestones, Germany PLOS ONE, 12 DOI Author Url Publisher Url Public Url

Davies TG, Rahman IA, Lautenschlager S, Cunningham JA, Asher RJ, Barrett PM, Bates KT, Bengston S, Benson RBJ, Boyer DM, Braga J, Bright JA, Claessens LPAM, Cox PG, Dong XP, Evans AR, Falkingham PL, Friedman M, Garwood RJ, Goswami A, Hutchinson JR, Jeffrey NS, Johanson Z, Lebrun R, Martínez-Pérez C, Marugán-Lobón J, O'Higgins PM, Metscher B, Orliac M, Rowe T, Rucklin M, Sánchez-Villagra MR, Shubin NH, Smith SY, Starck JM, Stringer C, Summers AP, Sutton MD, Walsh SA, Weisbecker V, Witmer LM, Wroe S, Yin Z, Rayfield E, Donoghue PCJ. 2017. Open data and digital morphology Proceedings of the Royal Society B: Biological Sciences, 284 DOI Author Url Publisher Url Public Url

Xing L, Lockley MG, Klein H, Falkingham PL, Kim JY, McRea RT, Zhang J, Persons IV WS, Wang T, Wang Z. 2016. First early Jurassic small ornithischian tracks from Yunnan Province, Southwestern China PALAIOS, 31 DOI Author Url Publisher Url Public Url

Falkingham PL, Horner AM. 2016. Trackways Produced by Lungfish During Terrestrial Locomotion Scientific Reports, 6 DOI Author Url Publisher Url Public Url

Klein H, Wizevich MC, Thuring B, Marty D, Thuring S, Falkingham PL, Meyer CA. 2016. Triassic chirotheriid footprints from the Swiss Alps: ichnotaxonomy and depositional environment (Cantons Wallis & Glarus) Swiss Journal of Palaeontology, 135 :295-314 DOI Author Url Publisher Url Public Url

Bates KT, Mannion PD, Falkingham PL, Brusatte S, Hutchinson JR, Otero A, Sellers WI, Sullivan C, Stevens K, Allen V. 2016. Temporal and phylogenetic evolution of the sauropod dinosaur body plan Royal Society Open Science, DOI Author Url Publisher Url Public Url

Xing L, Daqing L, Falkingham PL, Lockley M, Benton MJ, Klein H, Zhang J, Ran H, Persons IV WS, Dai H. 2016. Digit-only sauropod pes trackways from China – evidence of swimming or a preservational phenomenon? Scientific Reports, DOI Author Url Publisher Url Public Url

Basu C, Falkingham PL, Hutchinson JR. 2016. The extinct, giant giraffid Sivatherium giganteum – skeletal reconstruction and body mass estimation Biology Letters, 12 DOI Author Url Publisher Url Public Url

Rahman I, Zamora S, Falkingham PL, Phillips JC. 2015. Cambrian cinctan echinoderms shed light on feeding in the ancestral deuterostome Proceedings of the Royal Society B: Biological Sciences, 282 DOI Author Url Publisher Url Public Url

Bates KT, Falkingham PL, Macaulay S, Brassey C, Maidment SCR. 2015. Downsizing a giant: Re-evaluating Dreadnoughtus body mass Biology Letters, 11 :1-6 DOI Author Url Publisher Url Public Url

Falkingham PL, Gatesy SM. 2014. The birth of a dinosaur footprint: Subsurface 3D motion reconstruction and discrete element simulation reveal track ontogeny Proceedings of the National Academy of Sciences of the United States of America, 111 :18279-18284 DOI Author Url Publisher Url Public Url

Bennett MR, Morse SA, Falkingham PL. 2014. Tracks made by swimming Hippopotami: An example from Koobi Fora (Turkana Basin, Kenya) Palaeogeography, Palaeoclimatology, Palaeoecology, 409 :9-23 DOI Author Url Publisher Url Public Url

Maidment SCR, Bates KT, Falkingham PL, VanBuren C, Arbour V, Barrett PM. 2014. Locomotion in ornithischian dinosaurs: an assessment using three-dimensional computational modelling BIOLOGICAL REVIEWS, 89 :588-617 DOI Author Url Publisher Url

Falkingham PL, Bates KT, Farlow JO. 2014. Historical photogrammetry: Bird's Paluxy River dinosaur chase sequence digitally reconstructed as it was prior to excavation 70 years ago. PLoS One, 9 :1-5 DOI Author Url Publisher Url Public Url

Razzolini NL, Vila B, Castanera D, Falkingham PL, Barco JL, Canudo JI, Manning PL, Galobart A. 2014. Intra-trackway morphological variations due to substrate consistency: the El Frontal dinosaur tracksite (Lower Cretaceous, Spain). PLoS One, 9 DOI Author Url Publisher Url Public Url

Falkingham PL. 2014. Interpreting ecology and behaviour from the vertebrate fossil track record JOURNAL OF ZOOLOGY, 292 :222-228 DOI Author Url Publisher Url

Falkingham PL, Hage J, Bäker M. 2014. Mitigating the Goldilocks effect: the effects of different substrate models on track formation potential Royal Society Open Science, 1 :2-9 DOI Author Url Publisher Url Public Url

White MA, Falkingham PL, Cook AG, Hocknull SA, Elliott DA. 2013. Morphological comparisons of metacarpal I for Australovenator wintonensis and Rapator ornitholestoides: implications for their taxonomic relationships ALCHERINGA, 37 :435-441 DOI Author Url Publisher Url

Castanera D, Vila B, Razzolini NL, Falkingham PL, Canudo JI, Manning PL, Galobart A. 2013. Manus track preservation bias as a key factor for assessing trackmaker identity and quadrupedalism in basal ornithopods. PLoS One, 8 :1-13 DOI Author Url Publisher Url Public Url

Bates KT, Savage R, Pataky TC, Morse SA, Webster E, Falkingham PL, Ren L, Qian Z, Collins D, Bennett MR, McClymont J, Crompton RH. 2013. Does footprint depth correlate with foot motion and pressure? JOURNAL OF THE ROYAL SOCIETY INTERFACE, 10 DOI Author Url Publisher Url

Falkingham PL. 2013. Low cost 3D scanning using off-the-shelf video gaming peripherals Journal of Paleontological Techniques, 11 :1-9 Public Url

Bennett MR, Falkingham PL, Morse SA, Bates KT, Crompton RH. 2013. Preserving the Impossible: Conservation of Soft-Sediment Hominin Footprint Sites and Strategies for Three-Dimensional Digital Data Capture PLoS ONE, :e60755-e60755 DOI Author Url Publisher Url Public Url

Castanera D, Vila B, Razzolini NL, Falkingham PL, Canudo JI, Manning PL, Galobart À. 2013. Correction: Manus Track Preservation Bias as a Key Factor for Assessing Trackmaker Identity and Quadrupedalism in Basal Ornithopods. PloS one, 8 DOI Publisher Url

Sellers WI, Hepworth-Bell J, Falkingham PL, Bates KT, Brassey CA, Egerton VM, Manning PL. 2012. Minimum convex hull mass estimations of complete mounted skeletons BIOLOGY LETTERS, 8 :842-845 DOI Author Url Publisher Url

Bates KT, Falkingham PL. 2012. Estimating maximum bite performance in Tyrannosaurus rex using multi-body dynamics BIOLOGY LETTERS, 8 :660-664 DOI Author Url Publisher Url

Falkingham PL, Bates KT, Mannion PD. 2012. Temporal and palaeoenvironmental distribution of manus- and pes-dominated sauropod trackways JOURNAL OF THE GEOLOGICAL SOCIETY, 169 :365-370 DOI Author Url Publisher Url

Bates KT, Benson RBJ, Falkingham PL. 2012. A computational analysis of locomotor anatomy and body mass evolution in Allosauroidea (Dinosauria: Theropoda) PALEOBIOLOGY, 38 :486-507 DOI Author Url Publisher Url

Falkingham PL. 2012. Acquisition of high resolution three-dimensional models using free, open-source, photogrammetric software PALAEONTOLOGIA ELECTRONICA, 15 DOI Author Url Publisher Url

Falkingham PL, Bates KT, Margetts L, Manning PL. 2011. The 'Goldilocks' effect: preservation bias in vertebrate track assemblages JOURNAL OF THE ROYAL SOCIETY INTERFACE, 8 :1142-1154 DOI Author Url Publisher Url

Farlow JO, O'Brien M, Kuban GJ, Datillo BF, Bates KT, Falkingham PL, Pinuela L, Rose A, Freels A, Kumagai C, Libben C, Smith J, Whitcraft J. 2011. Dinosaur tracksites of the Paluxy River Valley (Glen Rose Formation, Lower Cretaceous), Dinosaur Valley State Park, Sommervell County, Texas Proceedings of the V International Symposium about Dinosaur Palaeontology and their Environmen, :41-69

Falkingham PL, Bates KT, Margetts L, Manning PL. 2011. Simulating sauropod manus-only trackway formation using finite-element analysis BIOLOGY LETTERS, 7 :142-145 DOI Author Url Publisher Url

Falkingham PL, Margetts L, Manning PL. 2010. FOSSIL VERTEBRATE TRACKS AS PALEOPENETROMETERS: CONFOUNDING EFFECTS OF FOOT MORPHOLOGY PALAIOS, 25 :356-360 DOI Author Url Publisher Url

Bates KT, Falkingham PL, Rarity F, Hodgetts D, Purslow A, Manning PL. 2010. APPLICATION OF HIGH-RESOLUTION LASER SCANNING AND PHOTOGRAMMETRIC TECHNIQUES TO DATA ACQUISITION, ANALYSIS AND INTERPRETATION IN PALAEONTOLOGY Mills JP, Barber DM, Miller PE, Newton I. PROCEEDINGS OF THE ISPRS COMMISSION V MID-TERM SYMPOSIUM CLOSE RANGE IMAGE MEASUREMENT TECHNIQUES, 38 :68-73 Author Url

Falkingham PL, Agenbroad LD, Thompson K, Manning PL. 2010. Bird Tracks at the Hot Springs Mammoth Site, South Dakota, USA ICHNOS-AN INTERNATIONAL JOURNAL FOR PLANT AND ANIMAL TRACES, 17 :34-39 DOI Author Url Publisher Url

Falkingham PL, Milán J, Manning PL. 2010. A crocodylian trace from the Lance Formation (Upper Cretaceous) of Wyoming New Mexico Museum of Natural History and Science, Bulletin, 51 :171-174

Manning PL, Margetts L, Johnson MR, Withers PJ, Sellers WI, Falkingham PL, Mummery PM, Barrett PM, Raymont DR. 2009. Biomechanics of Dromaeosaurid Dinosaur Claws: Application of X-Ray Microtomography, Nanoindentation, and Finite Element Analysis ANATOMICAL RECORD-ADVANCES IN INTEGRATIVE ANATOMY AND EVOLUTIONARY BIOLOGY, 292 :1397-1405 DOI Author Url Publisher Url

Falkingham PL. 2009. Walking with dinosaurs Planet Earth, :12-13

Bates KT, Falkingham PL, Breithaupt BH, Hodgetts D, Sellers WI, Manning PL. 2009. HOW BIG WAS 'BIG AL'? QUANTIFYING THE EFFECT OF SOFT TISSUE AND OSTEOLOGICAL UNKNOWNS ON MASS PREDICTIONS FOR ALLOSAURUS (DINOSAURIA: THEROPODA) PALAEONTOLOGIA ELECTRONICA, 12 Author Url

Falkingham PL, Margetts L, Smith IM, Manning PL. 2009. Reinterpretation of palmate and semi-palmate (webbed) fossil tracks; insights from finite element modelling PALAEOGEOGRAPHY PALAEOCLIMATOLOGY PALAEOECOLOGY, 271 :69-76 DOI Author Url Publisher Url

Bates KT, Falkingham PL, Hodgetts D, Farlow JO, Breithaupt BH, O'Brien M, Matthews N, Sellers WI, Manning PL. 2009. Digital imaging and public engagement in palaeontology Geology Today, 25 :134-139 DOI Publisher Url

Manning PL, Ott C, Falkingham PL. 2008. A probable tyrannosaurid track from the Hell Creek Formation (Upper Cretaceous), Montana, United States PALAIOS, 23 :645-647 DOI Author Url Publisher Url

Bates KT, Breithaupt BH, Falkingham PL, Matthews NA, Hodgetts D, Manning PL. 2008. Integrated LiDAR & photogrammetric documentation of the Red Gulch dinosaur tracksite (Wyoming , USA) Proceedings of teh 8th Conference on Fossil Resources, :101-103

Poster

Jackson N, Wakely D, Falkingham PL. 2021. 3D marker tracking with GoPros and XMALab: A low-cost 3D tracking set-up Society of Experimental Biology Annual Meeting Public Url

Conference publication

Hatala KG, Falkingham PL, Gatesy SM. 2021. A new ontogenetic framework for analyzing fossil hominin tracks AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, 174 :45-45 Author Url

Hatala KG, Falkingham PL, Megherhi S, Perry DA, Cheleden S, Novotny J, Laidlaw DH, Gatesy SM. 2020. Merging biplanar X-ray, 3-D animation, particle simulation, and virtual reality to understand 3-D foot dynamics on deformable substrates AMERICAN JOURNAL OF PHYSICAL ANTHROPOLOGY, 89th Annual Meeting of the American-Association-of-Physical-Anthropologists (AAPA) 171 :117-117 Author Url

Marek RD, Falkingham PL, Bates KT. 2019. The Surrogate Arm: Assessing Factors Influencing Vertebral Morphology in the Avian Cervical Spine JOURNAL OF MORPHOLOGY, 280 :S173-S174 Author Url

Falkingham PL, Turner M, Gatesy SM. 2019. Generating and Testing Hypotheses of Dinosaur Foot Motions Using 3D-Digitized Tracks and Large-Scale Granular Simulations JOURNAL OF MORPHOLOGY, 280 :S118-S118 Author Url

Turner ML, Novotny J, Falkingham PL, Laidlaw DH, Gatesy SM. 2019. Where Does Footprint Morphology Come from? Developing Virtual Reality Visualizations for Exploring Dinosaur Track Formation JOURNAL OF MORPHOLOGY, 280 :S70-S70 Author Url

Crandell KE, Howe R, Falkingham P. 2019. Using 3D-Printed Models to Explore the Functional Consequences of Avian Beak Morphology JOURNAL OF MORPHOLOGY, 280 :S56-S56 Author Url

Strickson EC, Wilkinson DM, Hutchinson JR, Falkingham PL. 2019. No Small Feet: Investigating Mass Distribution and Heteropody in the Past and Present JOURNAL OF MORPHOLOGY, 280 :S226-S226 Author Url

Crandell KE, Howe RO, Cannon C, Falkingham PL. 2019. A comparative analysis of the hydrodynamics of beak shape kingfishers INTEGRATIVE AND COMPARATIVE BIOLOGY, Annual Meeting of the Society-for-Integrative-and-Comparative-Biology (SICB) 59 :E44-E44 Author Url

Glasgow S, Troelsen P, Falkingham PL, Marek RD. 2019. Stretching Evolution: Regionalisation and Neck Elongation in Plesiosaurs Integrative and Comparative Biology, Annual Meeting of the Society-for-Integrative-and-Comparative-Biology (SICB) 59 Author Url Public Url

Gatesy SM, Turner ML, Falkingham PL. 2018. CT Imaging of Dinosaur Footprints: Hidden Topography and the Origin of Track Diversity INTEGRATIVE AND COMPARATIVE BIOLOGY, Annual Meeting of the Society-for-Integrative-and-Comparative-Biology (SICB) 58 :E72-E72 Author Url

Turner ML, Falkingham PL, Gatesy SM. 2018. Avian Subsurface Foot Kinematics on Deformable Substrates INTEGRATIVE AND COMPARATIVE BIOLOGY, Annual Meeting of the Society-for-Integrative-and-Comparative-Biology (SICB) 58 :E437-E437 Author Url

Farlow JO, Robinson NJ, Kumagai CJ, Paladino FV, Falkingham PL, Elsey RM, Martin AJ. 2018. Trackways of the American Crocodile (Crocodylus acutus) in Northwestern Costa Rica: Implications for Crocodylian Ichnology Ichnos, 25 :30-65 DOI Author Url Publisher Url Public Url

Falkingham PL, Gatesy SM. 2016. Using extant taxa to inform studies of fossil footprints

Farlow JO, Robinson NJ, Paladino FV, Falkingham PL, Martin AJ. 2016. Trackways of the American crocodile, northwestern Costa Rica: implications for crocodylian ichnology. 131st Annual Meeting, Indiana Academy of Science, Abstracts, 131st Annual Meeting, Indiana Academy of Science :57-57

Farlow JO, Bates KT, Bonem RM, Dattilo BF, Falkingham PL, Gildner R, Jacene J, Kuban GJ, Martin AJ, O'Brien M, Whitcraft J. 2015. DINOSAUR FOOTPRINTS FROM THE GLEN ROSE FORMATION (PALUXY RIVER, DINOSAUR VALLEY STATE PARK, SOMERVELL COUNTY, TEXAS) Noto C. Early- and Mid-Cretaceous Archosaur Localities of North-Central Texas, SVP 2015 Meeting Field Trip Guidebook, 75th Annual Meeting of the Society of Vertebrate Palaeontology :14-37 Publisher Url Public Url

Falkingham PL, Gatesy SM. 2015. The birth of a dinosaur track: sub-surface 3-D motion reconstruction and discrete element simulation reveal footprint ontogeny' INTEGRATIVE AND COMPARATIVE BIOLOGY, Annual Meeting of the Society-for-Integrative-and-Comparative-Biology (SICB) 55 :E54-E54 Author Url

Falkingham PL, Gatesy SM. 2012. RECONSTRUCTING LIMB KINEMATICS OF SMALL BIPEDAL DINOSAURS TRAVERSING SEMI-FLUID SUBSTRATES JOURNAL OF VERTEBRATE PALEONTOLOGY, 72nd Annual Meeting of the Society-of-Vertebrate-Paleontology 32 :92-92 Author Url

Falkingham P, Bates K, Mannion P. 2011. EVOLUTIONARY TRENDS IN SAUROPOD BODY PLAN EXPRESSED IN FOSSIL TRACKS AND TRACKWAYS JOURNAL OF VERTEBRATE PALEONTOLOGY, 31 :107-107 Author Url

Falkingham P. 2009. DIVERSITY IN FOSSIL VERTEBRATE TRACK ASSEMBLAGES: PRESERVATIONAL BIAS DUE TO SEDIMENT CONSISTENCY AND ANIMAL MASS JOURNAL OF VERTEBRATE PALEONTOLOGY, 29 :93A-93A Author Url

Falkingham P, Margetts L, Manning P. 2008. USING FINITE ELEMENT ANALYSIS TO AID INTERPRETATION OF DINOSAUR TRACKS JOURNAL OF VERTEBRATE PALEONTOLOGY, 68th Annual Meeting of the Society-of-Vertebrate-Paleontology 28 :76A-76A Author Url

Falkingham P, Manning P, Margetts L. 2007. Finite element analysis of dinosaur tracks JOURNAL OF VERTEBRATE PALEONTOLOGY, 27 :73A-73A Author Url

Farlow JO, Richards RL, Johnson BD, Falkingham PL. New fossil vertebrate specimens from the Pipe Creek Sinkhole (Grant County, Indiana) 130th Annual Meeting, Indiana Academy of Science, Program, 130th Annual Meeting, Indiana Academy of Science :38-38

Other

Bates KT, Falkingham PL. 2018. Estimating maximum bite performance in Tyrannosaurus rex using multi-body dynamics (vol 8, 20120056, 2012) BIOLOGY LETTERS, 14 DOI Author Url Publisher Url

Falkingham P. Fossil Bird Track, Mammoth Site of Hot Springs DOI

Falkingham P, Milan J. Emu track and Undertracks in cement - photos + photogrammetric models. DOI

Chapters

Falkingham PL. 2016. Applying objective methods to subjective track outlines. Falkingham PL, Marty D, Richter A. Dinosaur Tracks: The Next Steps :72-80 Indiana University Press 978-0-253-02102-1

Falkingham PL, Marty D, Richter A. 2016. Introduction Falkingham PL, Marty D, Richter A. Dinosaur Tracks The Next Steps :2-11 Indiana University Press 9780253021021

Marty D, Falkingham PL, Richter A. 2016. Dinosaur Track Terminology: A Glossary of Terms Falkingham PL, Marty D, Richter A. Dinosaur Tracks The Next Steps :399-402 Indiana University Press 9780253021021

Milan J, Falkingham PL. 2016. Experimental and Comparative ichnology Falkingham PL, Marty D, Richter A. Dinosaur Tracks: The Next Steps :14-27 Indiana University Press 978-0-253-02102-1

Milàn J, Falkingham PL. 2016. Experimental and comparative ichnology Dinosaur Tracks: The Next Steps :15-27 9780253021021

Manning PL, Falkingham PL. 2011. Science communication with dinosaurs Handbook of Research on Computational Science and Engineering: Theory and Practice :587-610 9781613501160 DOI Publisher Url

Manning PL, Falkingham PL. 2011. Science Communication with Dinosaurs Leng J, Sharrock W. Handbook of Research on Computational Science and Engineering: Theory and Practice :587-611 9781613501160 DOI

Books (authored)

Falkingham PL, Marty D, Richter A. 2016. Dinosaur Tracks The Next Steps Falkingham PL, Marty D, Richter A. 9780253021021

Falkingham PL, Marty D, Richter A. 2016. Introduction :3-11 9780253021021

Software/Code

Budka M, Bakirov R, Deng S, Falkingham PL, Reynolds SC, Bennett MR. 2016. DigTrace Academic

Budka M, Bakirov R, Deng S, Falkingham PL, Reynolds S, Bennett M. 2016. DigTrace Pro

Thesis/Dissertation

Falkingham PL. 2010. Commputer simulation of dinosaur tracks Manning P, Margetts L.

Media

Falkingham P, Xing L. Xing et al 2016 Anomoepus.zip DOI

Dataset

Hatala K, Gatesy S, Falkingham P. Integration of biplanar X-ray, 3-D animation, and particle simulation reveals details of human ‘track ontogeny’ - Data DOI

Farlow JO, Falkingham P. Fossil Screen-Wash set-up, Pipe Creek Sinkhole DOI

Farlow JO, Falkingham P. Canid Jaw - Pipecreek Sinkhole DOI

Falkingham P. Penarth tracks dataset DOI

Falkingham P, Horner AM. Lungfish terrestrial traces DOI

Highlighted activities

Research Grants Awarded:

UKRI, Mud to Muscles: Dinosaur to bird evolution from fossil footprints, Grant value (£): £2.2 million, Duration of research project: 5 years. 2023

Partnership for Advanced Computing in Europe, Simulating footprint formation, from Humans to Dinosaurs, Grant value (£): ~200,000 (15,000,000 CPU hours), Duration of research project: 12 months. 2022

Leverhulme, Bones in motion: A new integrated experimental-computational approach to understanding the evolution of human locomotion, Karl Bates, Kristian D'Aout, Grant value (£): 249045, Duration of research project: 3 years. 2017

NERC, Integrated software solution for the 3-dimensional capture and analysis of footwear evidence, Bennett; Budka; Gabrys; Osselton, Grant value (£): 100k, Duration of research project: 12 months. 2014

NSF, Birth of a dinosaur track: Formation dynamics and the origin of footprint diversity., Gatesy, S.M. (PI), Grant value (£): $315,698, Duration of research project: 3 years. 2014

NSF, X-rays, 3D animation and human locomotion, Kevin Hatala, Stephen Gatesy, Grant value (£): $340,052.

Professional activities

Public engagement:

Performance art: music, dance, drama, etc., Netflix viewers, Consultant/Biomechanics specialist, Consultant for Netflix Documentary "Life on Our Planet", Major consultant (credited) for netflix documentary on extinct life., https://www.netflix.com/gb/title/80213846. 2023

Other: Please specify in text box below, Global audience interested in palaeontology, Podcast, Guest, Podcast guest - Palaeocast, Interviewed for the podcast Palaeocast about my role consulting for a recent Netflix documentary, and the importance of biomechanics for media., https://www.palaeocast.com/life-on-our-planet/. 2023

Performance art: music, dance, drama, etc., TV watchers Channel 5/Amazon, Expert, Online/Terrestrial TV (Channel 5 and Amazon Prime), Documentary Appearance, Appeared on documentary "Dinosaurs with Stephen Fry", https://amzn.to/3sC260A. 2023

Public talk or lecture, Interest/students, Invited speaker, Earth Sciences department, Cambridge University, Talk for Sedgewick Society, University of Cambridge. 2019

Public talk or lecture, Undergraduate/interested public, Invited speaker, Bangor University, Talk for Bangor Biological Society, https://www.facebook.com/events/719496735076323. 2018

Public talk or lecture, 16-19 + parents, Invited speaker, Birkenhead Academy High School, Talk at Birkenhead Academy. 2018

Other: Please specify in text box below, General public, Online article, Author, https://theconversation.com/footprint-fossils-suggest-lizards-have-been-running-on-two-feet-for-110m-years-91865, Article for the Conversation: Footprint fossils suggest lizards have been running on two feet for 110m years. 2018

Other: Please specify in text box below, Wider Public, Article, Writer, https://theconversation.com/walking-fish-help-scientists-to-understand-how-we-left-the-ocean-91411, Article for The Conversation: ‘Walking’ fish help scientists to understand how we left the ocean. 2018

Public talk or lecture, Student society, Invited speaker, Bangor University, Talk for Bangor University Biology Socity. 2018

Other: Please specify in text box below, Global, Radio/New media, Expert, https://www.bbc.co.uk/programmes/w3cswvxr, BBC World Service Radio Programme - Crowd Science, https://www.bbc.co.uk/programmes/w3cswvxr. 2018

Other: Please specify in text box below, General; technical; specialist, Blog/Website, Author, https://pfalkingham.wordpress.com, Personal Website/Blog 2018. 2018

Other: Please specify in text box below, General/General science, Website/Blog, Writer/Author, https://pfalkingham.wordpress.com/, Personal Website/Blog 2017, https://pfalkingham.wordpress.com/. 2017

Other: Please specify in text box below, general public, Podcast, Interviewee, Palaeocast is a monthly podcast series, Online, Palaeocast interview, http://www.palaeocast.com/episode-44-trackway-modelling/. 2015

Other: Please specify in text box below, General Public, Online learning resource, Editor/Contributor, Online, Palaeontology[Online], http://www.palaeontologyonline.com/. 2011

Other: Please specify in text box below, general public/you, Online public outreach forum, Specialist, Online, AskABiologist Specialist, http://www.askabiologist.org.uk/. 2010

Research Grants Awarded:

UKRI, Mud to Muscles: Dinosaur to bird evolution from fossil footprints, Grant value (£): £2.2 million, Duration of research project: 5 years. 2023

Partnership for Advanced Computing in Europe, Simulating footprint formation, from Humans to Dinosaurs, Grant value (£): ~200,000 (15,000,000 CPU hours), Duration of research project: 12 months. 2022

Palaeontological Association, Regionalization of cervical vertebrae in plesiosaurs, Grant value (£): £2192, Duration of research project: 8 weeks. 2018

Leverhulme, Bones in motion: A new integrated experimental-computational approach to understanding the evolution of human locomotion, Karl Bates, Kristian D'Aout, Grant value (£): 249045, Duration of research project: 3 years. 2017

EPSRC/NERC, Extrapolating ground reaction force from fossil dinosaur footprints, Grant value (£): 63,334.90, Duration of research project: 24 months. 2015

Company of Biologists, Digitizing archosaur tracks and traces, Grant value (£): 500. 2015

NERC, Integrated software solution for the 3-dimensional capture and analysis of footwear evidence, Bennett; Budka; Gabrys; Osselton, Grant value (£): 100k, Duration of research project: 12 months. 2014

NSF, Birth of a dinosaur track: Formation dynamics and the origin of footprint diversity., Gatesy, S.M. (PI), Grant value (£): $315,698, Duration of research project: 3 years. 2014

XSEDE, USING COMPUTER SIMULATION TO STUDY DINOSAUR BIOMECHANICS FROM FOSSIL FOOTPRINTS (TG-EAR140012), S.M. Gatesy, Grant value (£): 365768 SUs, Duration of research project: 18 Months. 2014

NSF, X-rays, 3D animation and human locomotion, Kevin Hatala, Stephen Gatesy, Grant value (£): $340,052.

Media Coverage:

Featured on science show 'Quirks and Quarks' 2022
www.cbc.ca
www.cbc.ca

News coverage of Current Biology paper - Sauropod gaits 2022
institutions.newscientist.com
www.dailymail.co.uk
www.science.org

Dinosaur tracks found in south Wales 2022
www.thenational.wales
www.itv.com
www.mirror.co.uk

Media coverage associated with the Swimming Hippo Footprints paper published in Palaeo3 (Lead author: Bennett)

Daily Mail article on Lungfish traces

Blog post for the Gaurdian newspaper about the outreach site Palaeontology[Online]

Coverage on Dinosaur track CT work

Coverage of the announcement that the NERC grant (PI Bennett, Bournemouth University) has been awarded funding.
www.forensicmag.com
www1.bournemouth.ac.uk

Media coverage associate with the paper 'Historical Photogrammtery' publishing in PLOS ONE.
phenomena.nationalgeographic.com
www.bbc.co.uk
www.voanews.com

Media coverage for Bates et al 2016
www.engadget.com
phys.org
www.sciencedaily.com

Media coverage for paper 'Downsizing a Giant'
www.dailymail.co.uk
www.washingtonpost.com
news.nationalgeographic.com

Media Coverage for Sivatherium reconstruction
www.dailymail.co.uk
www.bbc.co.uk
www.nature.com

Media coverage of Digit-only sauropod tracks from China
www.bristol.ac.uk
www.csmonitor.com
www.asianscientist.com

Media coverage of PhD student (Pernille Troelsen) work on Plesiosaur necks.
knowridge.com
www.sciencedaily.com

Media coverage on the 2014 paper "The Birth of a Dinosaur Track" published in PNAS
sciencenode.org
www.bbc.co.uk
www.foxnews.com

News coverage on PLOS ONE ammonite paper
www.livescience.com
www.bbc.co.uk

Award:

Excellence in Digital Education, Teaching and Learning Academy, LJMU. 2022

Excellence in Digital Education Award Commendation. 2021

General Biomechanics session Talk Prize, Society for Experimental Biology. 2013

Conference presentation:

3D marker tracking with GoPros and XMALab: A low-cost 3D tracking set-up, Society of Experimental Biology, Poster presentation. 2021

WHY DID SOME QUADRUPEDAL DINOSAURS HAVE SMALL FRONT FEET?, SVP, Brisbane, Australia, Oral presentation. 2019

CT IMAGING OF DINOSAUR FOOTPRINTS: HIDDEN TOPOGRAPHY AND THE ORIGIN OF PENETRATIVE TRACK DIVERSITY, SVP, Brisbane, Australia, Oral presentation. 2019

Generating and testing hypothesis of dinosaur foot motions using 3d-digitized tracks and large-scale granular simulations., ICVM, Prague, Czech Republic, Oral presentation. 2019

No small feet: Investigating mass distribution and heteropody in the past and present., ICVM, Prague, Czech Republic., Oral presentation. 2019

Repeated evolution of drag reduction at the air-water interface in diving kingfishers, ICVM, Prague, Czech Republic., Oral presentation. 2019

The Surrogate Arm: Assessing factors influencing vertebral morphology in the avian cervical Spine., ICVM, Prague, Czech Republic, Oral presentation. 2019

Where does footprint morphology come from? Developing virtual reality visualizations for exploring dinosaur track formation, ICVM, Prague, Czech Republic., Oral presentation. 2019

A comparative analysis of the hydrodynamics of beak shape kingfishers, Society for Integrative and Comparative Biology Annual Meeting, Tampa, Florida, USA, Oral presentation. 2019

Stretching Evolution: Regionalisation and Neck Elongation in Plesiosaurs, Society for Integrative and Comparative Biology Annual meeting, Tampa, Florida, USA, Poster presentation. 2019

Developing Virtual Reality Visualizations of Dinosaur Track Creation with Scientific Skethcing, Vast Infovis SciVis (VIS) 2018, Berlin, Germany, Poster presentation. 2018

Where does footprint morphology come from? Integrating 3D methods for exploring dinosaur track formation, Society of Vertebrate Paleontology, Albuquerque, New Mexico, USA, Oral presentation. 2018

An everyday tool: how the avian neck has adapted to (almost) every situation., Symposium of Vertebrate Palaeontology and Comparative Anatomy, Manchester, UK, Oral presentation. 2018

Functional morphology and hydrodynamics of plesiosaur necks: Does size matter?, Symposium of Vertebrate Palaeontology and Comparative Anatomy, Manchester, UK, Oral presentation. 2018

The implications of underfoot pressures and mass distribution for extreme heteropody, Society of Experimental Biology, Florence, Italy, Poster presentation. 2018

The surrogate arm: Analysing the role of regionalisation in the variation of the avian neck, Society of Exoerimental Biology, Florence, Italy, Oral presentation. 2018

Avian Subsurface Foot Kinematics on Deformable Substrates, Society of Integrative and Comparative Biology, San Francisco, CA, USA, Poster presentation. 2018

CT Imaging of Dinosaur Footprints: Hidden Topography and the Origin of Track Diversity, Society of Integrative and Comparative Biology, San Franscisco, CA, USA, Oral presentation. 2018

REGIONALISATION OF THE AVIAN CERVICAL COLUMN: A LINK BETWEEN MORPHOLOGY AND ECOLOGY, Society of Vertebrate Paleontology, Calgary, Canada, Poster presentation. 2017

GRABBING EVOLUTION BY THE THROAT: FUNCTIONAL REGIONALISATION OF THE AVIAN CERVICAL COLUMN, Symposium on Vertebrate Palaeontology and Comparative Anatomy, Birmingham, Poster presentation. 2017

PHILOSOPHICAL PERSPECTIVES ON BIOMECHANICAL RECONSTRUCTIONS OF EXTINCT ANIMALS: A CASE STUDY USING TYRANNOSAURUS REX, Symposium on Vertebrate Palaeontology and Comparative Anatomy, Birmingham, Oral presentation. 2017

RANGE OF MOTION AND HYDRODYNAMIC IMPLICATIONS OF THE LONG- NECK IN PLESIOSAURS, Symposium on Vertebrate Palaeontology and Comparative Anatomy, Birmingham, Oral presentation. 2017

A re-analysis of the middle Pleistocene hominin-like footprints from Vértesszőlős, Hungary, British Association for Biological Anthropology and Osteoarchaeology, Liverpool, Poster presentation. 2017

A RE-ANALYSIS OF ~350 KA HOMININ-LIKE FOOTPRINTS FROM VÉRTESSZOLOS, HUNGARY, EMPLOYING PHOTOGRAMMETRY AND 3D ANALYSIS., Society of Vertebrate Paleontology, Calgary, Poster presentation. 2017

Can skeletal surface area predict in vivo foot surface area, Society of Experimental Biology, Sweden, Poster presentation. 2017

Grabbing evolution by the throat: Functional regionalisation of the avian cervical column, Society of Experimental Biology, Sweden, Poster presentation. 2017

Hydrodynamic implications of the long neck in plesiosaurs using computational fluid dynamics, Society of Experimental Biology, Gotenburg, Sweden, Poster presentation. 2017

Reconstructing moving morphology using RaspberryPi (PiROMM): Range of motion in ostrich cervical vertebrae at progressive stages of dissection, Society of Experimental Biology, Gotenburg, Sweden, Poster presentation. 2017

Foot Morphology or exit feature: A CT perspective on Hitchcock's Dinosaur Tracks, Symposium of Vertebrate Paleontology, Salt Lake City, UT, USA, Poster presentation. 2016

Footprints are neither bones nor feet: Problems with track "preservation" and the mold-based paradigm, Symposium of Vertebrate Paleontology, Salt Lake City, UT, USA, Poster presentation. 2016

The morphology of motion in sub-surface foot trajectories and fossil dinosaur tracks, Society of Vertebrate Paleontology Annual Meeting, Salt Lake City, UT, USA, Oral presentation. 2016

Trackways of the American Crocodile (Crocodylus Acutus), northwest Costa Rica: Implications for archosaurian ichnology, Society of Vertebrate Paleontology Annual Meeting, Salt Lake City, UT, USA, Poster presentation. 2016

Mobility and hydrodynamic implications for the long neck in plesiosaurs, Symposium of Vertebrate Palaeontology and Comparative Anatomy, Liverpool, Poster presentation. 2016

Making footprints without feet: Modern terrestrial lungfish locomotion creates traces similar to those of early tetrapods., Society of Experimental Biology (SEB), Brighton, UK, Oral presentation. 2016

One Foot, Many Footprints: the Origin of Track Morphological Diversity, International Conference on Vertebrate Morphology (ICVM), Washington DC, USA, Poster presentation. 2016

The morphology of Motion: Sub-surface foot trajectories and fossil tracks, International Congress on Vertebrate Morphology (ICVM), Washington DC, USA, Poster presentation. 2016

Using extant taxa to inform studies of fossil footprints, European Geosciences Union, Vienna, Austria, Oral presentation. 2016

Using LIGGGHTS to investigate dinosaur footprint formation, 1st CF-DEM User Meeting, Linz, Austria, Oral presentation. 2016

Difficulties in estimating mass from fossil footprints, SVPCA, Southampton, Oral presentation. 2015

Estimating body mass of the extinct giraffid Sivatherium giganteum, using a skeletal reconstruction and the minimum convex hull method, SVPCA, Southampton, Poster presentation. 2015

Conference organisation:

Symposium of Palaeontological Preparation and Conservation, Organiser/Host, http://svpca.org/. 2016

Symposium of Vertebrate Palaeontology and Comparative Anatomy, Host/Organiser. 2016

Progressive Palaeontology, http://www.palass.org/modules.php?name=propal&page=46. 2008

Other invited event:

European Geosciences Union Annual Meeting, Vienna, Austria, Invited speaker. 2016

Membership of professional bodies:

Member, European Geosciences Union. 2016

Member, Ichnological Association.

Member, Palaeontological Association.

Member, Society of Experimental Biology.

Member, Society of Integrative and Comparative Biology.

Member, Society of Vertebrate Paleontology.

Teaching qualification:

Fellow of the Higher Education Academy. 2015

Fellowships:

Marie Curie International Outgoing Fellowship, Royal Veterinary College, London. 2012

Editorial boards:

Journal of Paleontological Techniques, Scientific Editor, http://jpaleontologicaltechniques.org/. 2012

Palaeontology[Online], Commissioning Editor, http://www.palaeontologyonline.com/. 2011

External collaboration:

Brown University, Stephen Gatesy.

Liverpool University, Karl Bates.

Royal Veterinary College, John Hutchinson.

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